One inclusion formed somewhat later exterior to the exosporium. Another inclusion was formed within the exosporium and remained with the spore after mother cell lysis.

当芽成细裂解后，一种被释放出来，成自由的伴，另一种在芽内侧成，成熟后被包裹在芽内不脱落，始终和芽粘连在一起。